Carolinea 74

L amos : Tokukobelba gen. nov. (Acari: Oribatida: Damaeidae) 81 of priority. The publication of T rägårdh (1902) was printed on 7 th May of that year. K ulczy ń ski in- troduced his new species in not one, but two sep- arate papers published shortly after each other. One of these ( K ulczy ń ski 1902a) is written only in the Latin language, while in the other ( K ulczy ń ski 1902b) the text is partially in Latin and partially in German. I have not been able to determine the exact date of publication of the former, but a copy of the latter ( K ulczy ń ski 1902b) residing in the library of the Museum of Comparative Zool- ogy at Harvard University bears a library stamp of 28 April 1902. Hence T . farinosa is very likely a junior synonym of T compta and not the other way round. In the present publication only T . farinosa and T. verrucosa verrucosa are accepted as poten- tial synonyms of T compta . The subspecies T verrucosa japonica from Japan, introduced by A oki (1984), differs considerably from T verru­ cosa and is here treated as a full species. Based on the paper of A oki (1984), T japonica differs from T compta in: 1) smaller body size with a body length of 342-360 µm instead of 460-560 µm for European and Russian specimens of T compta ; 2) distinctly longer sensillus; 3) rostrum broadly rounded in dorsal perspective instead of narrow, pointed; 4) femora I and II with 5 and 4 setae, respectively; 5) associated seta d ab- sent on tibia IV. There are several discrepancies between the written description and the illus- trations of the species supplied. For example, A oki (1984, p. 111) states that the femur and tibia of leg I both possess 4 setae, whereas he (p. 113, Fig. 6c) depicts 5 setae on both of these segments. A oki (1984) describes his spe- cies as having only 9 pairs of notogastral setae. However, the pseudanal setae ps2 and ps3 are somewhat cryptic in Tokukobelba species since they are short and lie close to the notogastral surface and are hence easy to miss. The as- sociated seta d of tibia IV is very short and re- gressed in other Tokukobelba species. Possibly the protecting seta d was overlooked, but it may also have been strongly reduced or even lost in this species as was noted by B ayartogtokh (2000). Tokukobelba japonica needs to be thor- oughly redescribed. At present there is no solid basis for establishing a synonymy between this species and T compta . Species which may belong to or show affini- ties to Tokukobelba . There appear to be several as yet undescribed species within the new ge- nus. For example, the leg setation of T sellnicki from Russia ( B ulanova - Z achvatkina 1962) differs significantly from those of China as noted by W ang & N orton (1995), with the Chinese speci- mens being incompletely described. In a similar fashion, representatives of this species from Ko- rea ( C hoi & N amkoong 2002), for which the leg setation and ventral features are unknown, dif- fer from the Russian ones in their much shorter notogastral setae and a non flagellate sensillus. It is very likely that a detailed revision also incor- porating molecular data will increase the species count of the genus. A new species of Parabelbella B ulanova -Z ach - vatkina , 1975, was recently reported from China: Parabelbella dimidiaspina X ie , Y an & Y ang , 2013. Based on its original description this shares with Tokukobelba : notogastral setae c1 and c2 inserted very close together and directed poste- riad; sensillus barbed in the middle region, dis- tally flagellate; bothridia oriented distinctly pos- teriolaterad; apophysis Ba present with Bp, Da, Dp absent; centrodorsal apophyses Aa and Ap present; presence of numerous small tubercles on epimeral regions; solenidion of tibia III-IV elongate tactile. X ie et al. (2013, p. 280, Figs 6, 11-12) depict the tarsus of leg I with setae (v1) and (v2) both present, and with a pair of ventral setae located on each of tarsi III and IV. Tarsus I therefore very likely possesses 22 setae instead of the 20 given by these authors, and according- ly, the proral setae are not depicted in the illustra- tion of leg IV ( X ie et al. 2013, p. 280, Fig. 6). The authors state that the species displays an asso- ciated seta d on tibia II but do not illustrate this, while an associated seta d is instead depicted on tibia III (Fig. 10). The seta d here is not shown as sharing an alveolus with its solenidion φ , but neither is any of the genual setae d shown as inserting in the same alveolus as their associat- ed solenidia σ . Parabelbella dimidiaspina seems to be no Parabelbella but instead is an unusual Dyobelba N orton , 1979, with possibly very close affinities to Tokukobelba . This interesting species deserves detailed re-examination. The enigmatic, incompletely described species Costeremus yezoensis F ujikawa & F ujita , 1985, and C barbatus C ho i, 1997, both displaying a distinct damaeid habitus and located in the Hungarobelbidae M iko & T ravé , 1996, by S ubías (2004) and B ayartogtokh & E rmilov (2015), but in the Damaeolidae G randjean , 1965, by F ujikawa & F ujita (1985), C hoi (1997a) and C hen et al. (2010) instead, share several character states